Tricleidus

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Skull of Tricleidus in lateral view. From Brown (1981).

Tricleidus is a cryptoclidid from the Oxford Clay Formation of the UK. The holotype specimen (NHMUK R 3539) consists of disarticulated elements including most of skull and half the postcranium, from the Kosmoceras jasoni – Peltoceras athleta zones from the lowest deposits of the Oxford Clay Formation. The type and only species of Tricleidus is T. seeleyi. Persson (1962) erected...

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Thalassiodracon

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Thalassiodracon lectotype specimen (NHMUK 2018*).

The species Plesiosaurus hawkinsii was introduced in 1838 for a small plesiosaurian from Street, Somerset. The new genus name Thalassiodracon was erected decades later following an examination of a referred skull in Cambridge (CAMSM J.46986). Thalassiodracon is from the Rhaetian-Hettangian boundary, so it is latest Triassic or earliest Jurassic in age. Thalassiodracon means ‘Sea Dragon’, which “alludes to the colloquial...

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Simolestes

Simolestes has variously been allied with the Pliosauridae and the Rhomaleosauridae. The most noticeable difference between Simolestes and the other pliosaur taxa from the Oxford Clay (Liopleurodon, Peloneustes, Pachycostasaurus), is its much shorter snout and mandibular symphysis, a character is shares with the Rhomaleosauridae. However, this is probably a convergent character. Older descriptions of this taxon note the spatulate symphysis...

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Rhomaleosaurus

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Reconstruction of Rhomaleosaurus thorntoni. From Smith and Benson (2014).

Rhomaleosaurus is the largest known Lower Jurassic pliosaur and was the top predator in early Jurassic marine ecosystems. It has a reinforced skull to help resist torsion and a ferocious set of teeth, a combination of characters perfect for snatching and killing cephalopods, fish, and other marine reptiles. Historically, the genus Rhomaleosaurus has been interchangeable with now invalid ‘Thaumatosaurus’. See...

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Plesiosaurus

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Reconstruction of Plesiosaurus dolichodeirus in lateral view. From Conybeare (1824).

Plesiosaurus was the first plesiosaur discovered and named (De la Beche and Conybeare 1821). The first 1821 description was based on partial remains so the anatomy of Plesiosaurus – particularly the identity of the isolated bones and how they articulated together – involved some guesswork. An attempted reconstruction of the forelimb of Plesiosaurus by De la Beche and Conybeare (1821)....

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Peloneustes

Peloneustes is one of several pliosaur genera from the Oxford Clay Formation in the UK. It is one of the better known pliosaurs from this horizon, represented by 12 nearly complete skulls and several skeletons. It is also the most abundant pliosaur from the Peterborough Member of the Oxford Clay Formation (Ketchum and Benson 2011b). Peloneustes is characterised by an...

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Pachycostasaurus

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Mounted skeleton of Pachycostasaurus in the Peterborough Museum. Photo by Adam Smith.

Pachycostasaurus is approximately 3.1 meters long. Its rib cage and vertebrae exhibit thickened bone (Cruickshank et al. 1996) a condition termed pachyostosis. This heavy ossification is unusual in plesiosaurs (another exception may include Kronosaurus), although it is common in basal sauropterygians, especially pachypleurosaurs. Pachycostasaurus probably used the extra ballast provided by its heavy bones to traverse the sea bed searching...

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Macroplata

Macroplata is a rhomaleosaurid plesiosaur from the Lower Jurassic of Warwickshire, UK. Macroplata was named and briefly described by Swinton (1930a) based on a single skeleton from Harbury, Warwickshire, and the skeleton was redescribed in detail by Ketchum and Smith (2010). Photograph of Macroplata tenuiceps in situ in Harbury, Warwickshire. White (1940) referred the species Plesiosaurus longirostris to Macroplata, and...

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Liopleurodon

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Mounted skeleton of Liopleurodon ferox in the Tubingen Museum, Germany (from Martill and Naish, 2000).

Liopleurodon is a pliosaur that hardly needs introduction since appearing as the villain in the BBC’s ‘Walking with Dinosaurs’ TV series. This led to popular misconceptions about the size of Liopleurodon, which is known to have reached adult sizes in the region of seven metres long, nowhere near the gargantuan 25m estimate proposed by the series and later perpetuated elsewhere....

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Leptocleidus

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Skull of Leptocleidus capensis in lateral view (from Cruickshank, 1997)

Lower Cretaceous plesiosaurs are rare, so Leptocleidus is important because it fills a gap in the fossil record of plesiosaurians. Leptocleidus was once considered to be a late surviving member of the family Rhomaleosauridae but it has recently been reidentified as a close relative of polycotylids. The fossils of all known species of Leptocleidus were preserved in inshore brackish or...

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Kimmerosaurus

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Skull of Kimmerosaurus. From Brown (1981).

Kimmerosaurus is a poorly known cryptoclidid plesiosaur known only from skull and neck material (Brown 1981, Brown et al. 1986). It differs from other plesiosaurs in the large number of needle-like teeth in its jaws, and in the form of its teeth, which are completely smooth and lack the longitidinal ridges present in other plesiosaurs. Brown et al. (1986) compared...

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Eretmosaurus

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Skeleton of Eretmosaurus. Lithograph from Owen (1865).

The genus Eretmosaurus was erected for ‘Plesiosaurus’ rugosus by Seeley (1874). Eretmosaurus is a rogue taxon in cladistic analyses and researchers have been in disagreement about its taxonomic affinity. Eretmosaurus has been included in several different families: the Rhomaleosauridae based on the anatomy of its girdles (Persson 1963); the Pliosauridae (Brown 1981), the Elasmosauridae (Bardet 1995, Bardet et al. 1999). The...

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Cryptoclidus

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Skeleton of Cryptoclidus in lateral view. From Brown (1981).

Cryptoclidus, often wrongly spelled ‘Cryptocleidus’ after Andrews (1909), is a moderately sized plesiosaur with adults about four metres long (Brown 1981). It is known from a large number of individual specimens from the Oxford Clay Formation. Fossils of Cryptoclidus are relatively common, and provide a complete ontogenetic sequence from very young to old adult individuals. This makes Cryptoclidus one of...

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Atychodracon

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Holotype specimen of Atychodracon megacephalus in the Bristol City Museum and Art Gallery (from Swinton, 1948). The specimen was destroyed during the Second World War.

The genus Atychodracon was erected by Smith (2015) to accommodate ‘Rhomaleosaurus’ megacephalus, because it is generically separarate from Rhomaleosaurus sensu stricto (Smith and Dyke 2008). A. megacephalus is closely related to Eurycleidus and some authors have regarded A. megacephalus as a distinct species of Eurycleidus. Three dimensional scan with texture (colour) removed of plaster cast (BGS GSM 118410) of the...

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Attenborosaurus

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Photograph of the holotype of Attenborosaurus, before it was destroyed. The skeleton was mounted in a block together with a cast of the opposite side of the body and the opposite side of the head (from Swinton, 1949)

Attenborosaurus is an unusual plesiosaur because it combines a long neck with a relatively large head. It is classified as a pliosaur in some classifications, but some phylogenies place it in a more basal position. The genus was introduced for ‘Plesiosaurus’ conybeari, a species originally described by Sollas (1881). Bob Bakker coined the new genus name to honour the esteemed...

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