Pliosauroidea

The Pliosauroidea, or pliosaurs, are a superfamily of typically short-necked plesiosaurs.

Referred specimen of Peloneustes in the Tubingen Museum, Germany.

They usually have a low number of short vertebrae in the neck and often have huge skulls. In some genera the skull may reach up to 3 meters in length and constitute about a quarter of the total length of the animal. The clade includes two main families Pliosauridae and Rhomaleosauridae (although some recent phylogenies place the latter outside of Pliosauridae in a more basal position) as well as a variety of basal genera whose position is uncertain. Some authors also recognise a derived family of pliosaurs called the Brachauchenidae, which includes Brachauchenius and Kronosaurus.

Pliosauroid genera

Anguanax

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Anningasaura

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Archaeonectrus

The genus Archaeonectrus was proposed by Novozhilov (1964) for ‘Plesiosaurus’ rostratus, a species named by Owen (1865). By modern standards, Owen’s (1865) original description is rather inadequate. One notable characteristic of Archaeonectrus rostratus is the relatively small size of the limbs relative to its body. It has been classified as a pliosauroid, sometimes within the …

Attenborosaurus

Attenborosaurus is an unusual plesiosaur because it combines a long neck with a relatively large head. It is classified as a pliosaur in some classifications, but some phylogenies place it in a more basal position. The genus was introduced for ‘Plesiosaurus’ conybeari, a species originally described by Sollas (1881). Bob Bakker coined the new genus …

Atychodracon

The genus Atychodracon was erected by Smith (2015) to accommodate ‘Rhomaleosaurus’ megacephalus, because it is generically separarate from Rhomaleosaurus sensu stricto (Smith and Dyke, 2008). A. megacephalus is closely related to Eurycleidus and some authors have regarded A. megacephalus a distinct species of Eurycleidus. The holotype specimen of A. megacephalus was housed in the Bristol …

Bishanopliosaurus

Under construction

Borealonectes

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Cryonectes

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Eurycleidus

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Gallardosaurus

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Hauffiosaurus

Hauffiosaurus is a basal pliosaurid known from the Lower Jurassic (Toarcian) of the UK and Germany. Hauffiosaurus is a medium to large sized plesiosaur (H. zanoni = 3.4 m, H. tomistomimus, 4.23 m, H. longirostris = 4.83 m) (Smith and Lomax, 2019), with a relatively long neck for a pliosaur, and an elongate narrow snout. …

Kronosaurus

Due to its large size and ferocious appearance, Kronosaurus is one of the most famous plesiosaurs. The iconic skeleton referred to Kronosaurus on display in the Museum of Comparative Zoology in Harvard is nicknamed ‘plasterosaurus’ because so much of it is reconstructed in plaster. About a third of the skeleton is plaster and there are …

Liopleurodon

Liopleurodon is a pliosaur that hardly needs introduction since appearing as the villain in the BBC’s ‘Walking with Dinosaurs’ TV series. This led to popular misconceptions about the size of Liopleurodon, which is known to have reached adult sizes in the region of seven metres long, nowhere near the gargantuan 25m estimate proposed by the …

Maresaurus

M. coccai (Gasparini, 1997) type material: MOZ 4386 V articulated skull and mandible, atlas-axis and first cervical vertebrae. From the upper part of the Los Molles Formation, Cuyo Group, Emileia giebeli and Emileia multiformis subzone.

Marmornectes

Under construction

Megalneusaurus

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Meyerasaurus

M. victor was originally described and figured by Fraas (1910). Historically, M. victor has been regarded as a species of Rhomaleosaurus and is often associated in the literature under the defunct name ‘Thaumatosaurus’. Smith and Vincent (2010) identified M. victor as generically distinct from Rhomaleosaurus and erected the new name Meyerasaurus for it (Smith and Dyke, 2008). The …

Pachycostasaurus

Pachycostasaurus is approximately 3.1 meters long. Its rib cage and vertebrae exhibit thickened bone (Cruickshank et al. 1996) a condition termed pachyostosis. This heavy ossification is unusual in plesiosaurs (another exception may include Kronosaurus), although it is common in basal sauropterygians, especially pachypleurosaurs. Pachycostasaurus probably used the extra ballast provided by its heavy bones to …

Peloneustes

Peloneustes is one of several pliosaur genera from the Oxford Clay Formation in the UK. It is one of the better known pliosaurs from this horizon, represented by 12 nearly complete skulls and several skeletons. It is also the most abundant pliosaur from the Peterborough Member of the Oxford Clay Formation (Ketchum and Benson, 2011). …

Pliosaurus

Under construction

Rhomaleosaurus

Rhomaleosaurus is the largest known Lower Jurassic pliosaur and was the top predator in early Jurassic marine ecosystems. It has a reinforced skull to help resist torsion and a ferocious set of teeth, a combination of characters perfect for snatching and killing cephalopods, fish, and other marine reptiles. Historically, the genus Rhomaleosaurus has been interchangeable …

Simolestes

Simolestes has variously been allied with the Pliosauridae and the Rhomaleosauridae. The most noticeable difference between Simolestes and the other pliosaur taxa from the Oxford Clay (Liopleurodon, Peloneustes, Pachycostasaurus), is its much shorter snout and mandibular symphysis, a character is shares with the Rhomaleosauridae. However, this is probably a convergent character. Older descriptions of this …

Stenorhynchosaurus

Under construction

Thalassiodracon

The species Plesiosaurus hawkinsii was introduced in 1838 for a small plesiosaurian from Street, Somerset. The new genus name Thalassiodracon was erected decades later following an examination of a referred skull in Cambridge (CAMSM J.46986). Thalassiodracon means ‘Sea Dragon’, which “alludes to the colloquial description given to the Street marine reptile fauna by Hawkins” (Storrs …

Thaumatodracon

Thaumatodracon is a relatively large rhomaleosaurid from the Lower Jurassic (Sinemurian) of Lyme Bay ā€“ the coast between Lyme Regis and Charmouth ā€“ UK. The holotype specimen (NLMH 106.058) is an almost complete skull and cervical (neck) series. It has a 60 cm long skull, and based on comparison with other rhomaleosaurids I estimate its total …