Plesiosauroidea

Abyssosaurus is a derived cryptoclidid plesiosaur from the Upper Hauterivian (Lower Cretaceous) of the Menya River, Chuvashia, Russia. It was named and described in 2011 by Alexander Yu Berezin (Berezin 2011). A partial skull associated with the holotype specimen (MChEIO, no.

Albertonectes is a very long-necked elasmosaurid from the Late Cretaceous of Alberta, Canada. The holotype specimen (TMP 2007.0110001) consists of an almost complete skeleton lacking a skull (Kubo et al. 2012). The neck contains 76 cervical vertebrae, which is a unique character of Albertonectes vanderveldei and the highest number of neck vertebrae known for any plesiosaur, surpassing the previous record-holder (Elasmosaurus) by four vertebrae.

The genus and species ‘Apractocleidus teretipes‘ was erected by Smellie (1916) for a specimen now regarded as old-adult individual of Cryptoclidus. The specimen was collected by Alfred Leeds and acquired by the Hunterian Museum, Glasgow.

The mysterious plesiosaur Aristonectes is notable for its mouthful of pin-like teeth. A special feeding guild, the ‘trap guild’, has been proposed to accommodate Aristonectes and other plesiosaurs with similar dentition (Chatterjee and Small 1989) such as Cryptoclidus, Kimmerosaurus, and Kaiwhekea.

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Skeleton of Brancasaurus in side view. From Sachs et al. (2016)

Skull of Brancasaurus in dorsal and lateral view, and showing the tip of the snout (C) in ventral view (from Wegner 1914)

Colymbosaurus is a colymbosaurine (derived cryptoclidid) from the Late Jurassic of the UK and Svalbard, Norway. Two valid species are known, C. megadeirus from the UK, and C. svalbardensis from Svalbard, Norway.

Cryptoclidus, often wrongly spelled ‘Cryptocleidus’ after Andrews (1909), is a moderately sized plesiosaur up to 3 metres long. It is known from a large number of individual specimens from the Oxford Clay Formation. Fossils of Cryptoclidus are relatively common, and provide a complete ontogenetic sequence from very young to old adult individuals.

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Edgarosaurus is a basal polycotylid (Druckenmiller 2002) from the Thermopolis Shale

Elasmosaurus is one of the most widely recognised plesiosaur names and has become a stereotype for all elasmosaurids. However, it is relatively poorly known. The type and only known specimen of Elasmosaurus platyurus (ANSP 10081) includes the tip of the snout, occipital condyle, and the majority of the vertebral column.

‘Gronausaurus‘ was named in 2013 (Hampe 2013). It is now regarded as a junior synonym of Brancasaurus (Sachs et al. 2016).

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Hydrotherosaurus is a long-necked elasmosaurid from California, USA. It is represented by one of the most complete elasmosaurid skeletons ever discovered, so Hydrotherosaurus is one of the best known members of this family. The almost complete type skeleton of Hydrotherosaurus was discovered in the Panoche Hills by Mr.

Lower Cretaceous plesiosaurs are rare, so Leptocleidus is important because it fills a gap in the fossil record of plesiosaurians. Leptocleidus was once considered to be a late surviving member of the family Rhomaleosauridae but it has recently been reidentified as a close relative of polycotylids.

Libonectes was erected for ‘Elasmosaurus morgani’. The pectoral girdle and a forelimb of the holotype are figured (most recently by Welles 1962, fig. 12) but now lost (Carpenter 1999). Libonectes morgani (Welles 1949) is the type and only species. 

Lusonectes is a poorly known microcleidid the Toarcian (Lower Jurassic) of Portugal. It is known from a single holotype specimen (MG33), a partial skull in the Museu Geológico, Lisbon, Portugal. It was described and named by Smith et al.

Microcleidus is a medium (4.27 m) to large (5.03 m) plesiosauroid with a small head and a long neck containing 38–39 cervical vertebrae.

Watson (1909) erected Microcleidus to accommodate fossil material belonging to ‘P.’ homalospondylus and ‘P.’ macropterus (Watson 1911).

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Restoration of the skeleton of Muraenosaurus in lateral view. From Andrews 1910.

Posterior cervical (neck) vertebra of Muraenosaurus in lateral and posterior view. From Andrews (1910).

Dorsal vertebra of Muraenosaurus in anterior view.

Nakonanectes is a small elasmosaurid plesiosaur from the Bearpaw Shale of Montana, USA. It is known from a single moderately complete specimen including a particularly fine skull. It has a relatively short neck for an elasmosaur consisting of ‘only’ 39-42 neck vertebrae.

‘Occitanosaurus’ was described by Sciau et al. (1990) and Bardet et al. (1999). The genus name was erected for ‘Plesiosaurus’ tournemirensis by Bardet et al. (1999). The taxon was originally regarded as an early elasmosaurid but is now considered a member of the Microcleididae.

The type skeleton of ‘Ogmodirus’ was collected in 1909 by C. Boyce from the upper Greenhorn Limestone Formation (Lower Turonian, Late Cretaceous) of Cloud County (near Aurora), Kansas (Storrs 1999, Schumacher and Everhart 2005). The specimen, KUVP 441, is a partial skeleton consisting of partial vertebral column (51 cervical vertebrae, 18 caudal vertebrae), limb, and girdle elements.

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Plesiosaurus was the first plesiosaur discovered and named (De la Beche and Conybeare 1821). The first 1821 description was based on partial remains so the anatomy of Plesiosaurus – particularly the identity of the isolated bones and how they articulated together – involved some guesswork.

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The holotype specimen (KUVP 1301) of Styxosaurus is an articulated skull and anterior portion of the neck. It was found in the Niobrara Chalk near Hell Creek, Logan County, Western Kansas in 1890. It was described later that year as a new species Cimoliosaurus snowii by Williston (1890a) (and Williston 1890b), who subsequently (Williston 1906) referred the species to Elasmosaurus.

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The holotype of Thalassomedon was discovered by R. L. Landberg in 1939 in Baca County, Colorado. Thalassomedon has a short and deep atlas-axis and 62 cervical vertebrae, and there is no pectoral or pelvic bar in the adult condition (Carpenter 1999).

Thililua has 30 cervical vertebrae. Each cervical has a longitudinal ridge on its lateral surfaces, this is a convergent feature with elasmosaurids. The skull of Thililua is relatively large with a short postorbital region and elongate rostrum.

Tricleidus is a cryptoclidid from the Oxford Clay Formation of the UK. The holotype specimen (NHMUK R 3539) consists of disarticulated elements including most of skull and half the postcranium, from the Kosmoceras jasoni – Peltoceras athleta zones from the lowest deposits of the Oxford Clay Formation.

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Vectocleidus material on display in the Dinosaur Isle Museum, Isle of Wight, UK.

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Skeleton of Westphaliasaurus. The central slab of the skeleton was awaiting preparation at the time of this photograph, but was subsequently prepared. From Schwermann & Sander (2011).

Sachs (2004) regarded ‘Woolungasaurus‘ as a junior synonym of Styxosaurus and assigned the type species (‘W. glendowerensis‘) to that genus under the new combination ‘Styxosaurus glendoweresnis‘.