Pliosauroids, or colloquially just ‘pliosaurs’, are a superfamily of typically short-necked plesiosaurs. They usually have a small number of short neck vertebrae and often have huge skulls, but the earliest pliosaurs have a more conservative (in plesiosaur terms) body plan with a reasonably long neck and small head. In some pliosaur genera the skull may reach up to 3 meters long and make up about a quarter of the total length of the animal.
Pliosauroid cladogram
The clade traditionally includes two main families, the Pliosauridae and Rhomaleosauridae, as well as a small number of basal genera whose position is uncertain. Some phylogenies have placed the Rhomaleosauridae in a more basal position, outside of the Pliosauroidea (e.g. Benson et al. 2012, Benson and Druckenmiller 2014), so the position of Rhomaleosauridae within the plesiosaur family tree is uncertain. A clade of derived large-headed forms within the Pliosauridae is called Thalassophonea, and some authors also recognise a derived subfamily of thalassophonean pliosaurids called the Brachaucheninae (sometimes regarded as a family Brachauchenidae), which includes Brachauchenius and Kronosaurus.
Pliosauroid genera
Anguanax
Anguanax is the first articulated plesiosaur ever found in Italy. It is a basal thalassophonean pliosaur from the Late Jurassic (middle Oxfordian). The type and only known specimen (MPPL 18797) of Anguanax was originally described as an indeterminate pliosaurid (Cau and Fanti 2014), but was subsequently named after reanalysis by the same authors the following […]
Anningasaura
Anningasaura is a basal plesiosaur from the Lower Jurassic of Lyme Regis, UK, known from a single three-dimensionally preserved skull. The specimen, NHMUK OR49202, was originally described and figured by Andrews (1896), who referred it to ‘Plesiosaurus‘ macrocephalus. It was redescribed and named as the holotype of Anningasaura lymense by Vincent and Benson (2012). The […]
Archaeonectrus
The genus Archaeonectrus was proposed by Novozhilov (1964) for ‘Plesiosaurus’ rostratus, a species named by Owen (1865). By modern standards, Owen’s (1865) original description is rather inadequate. One notable characteristic of Archaeonectrus rostratus is the relatively small size of the limbs relative to its body. It has been classified as a pliosauroid, sometimes within the […]
Attenborosaurus
Attenborosaurus is an unusual plesiosaur because it combines a long neck with a relatively large head. It is classified as a pliosaur in some classifications, but some phylogenies place it in a more basal position. The genus was introduced for ‘Plesiosaurus’ conybeari, a species originally described by Sollas (1881). Bob Bakker coined the new genus […]
Atychodracon
The genus Atychodracon was erected by Smith (2015) to accommodate ‘Rhomaleosaurus’ megacephalus, because it is generically separarate from Rhomaleosaurus sensu stricto (Smith and Dyke 2008). A. megacephalus is closely related to Eurycleidus and some authors have regarded A. megacephalus as a distinct species of Eurycleidus. The holotype specimen of A. megacephalus (BRSMG Cb 2335) was […]
Bishanopliosaurus
Bishanopliosaurus is the most complete plesiosaur known from the Jurassic of Asia (Sato et al. 2003). The holotype specimen of the type species, B. youngi, is a partial postcranial skeleton of a juvenile individual from the Dongyuemiao Member of the Ziliujing Formation (Lower or Mid Jurassic) of Bishan County, Chongqing, China. It was first described […]
Borealonectes
Under construction
Cryonectes
Under construction
Eurycleidus
Under construction
Gallardosaurus
Under construction
Hauffiosaurus
Hauffiosaurus is a basal pliosaurid known from the Lower Jurassic (Toarcian) of the UK and Germany. Hauffiosaurus is a medium to large sized plesiosaur (H. zanoni = 3.4 m, H. tomistomimus, 4.23 m, H. longirostris = 4.83 m) (Smith and Lomax 2019), with a relatively long neck for a pliosaur, and an elongate narrow snout. […]
Kronosaurus
Due to its large size and ferocious appearance, Kronosaurus is one of the most famous plesiosaurs. The iconic skeleton referred to Kronosaurus on display in the Museum of Comparative Zoology in Harvard is nicknamed ‘plasterosaurus’ because so much of it is reconstructed in plaster. About a third of the skeleton is plaster and there are […]
Liopleurodon
Liopleurodon is a pliosaur that hardly needs introduction since appearing as the villain in the BBC’s ‘Walking with Dinosaurs’ TV series. This led to popular misconceptions about the size of Liopleurodon, which is known to have reached adult sizes in the region of seven metres long, nowhere near the gargantuan 25m estimate proposed by the […]
Macroplata
Macroplata is a rhomaleosaurid plesiosaur from the Lower Jurassic of Warwickshire, UK. Macroplata was named and briefly described by Swinton (1930a) based on a single skeleton from Harbury, Warwickshire, and the skeleton was redescribed in detail by Ketchum and Smith (2010). White (1940) referred the species Plesiosaurus longirostris to Macroplata, and this species has been […]
Maresaurus
M. coccai (Gasparini 1997) type material: MOZ 4386 V articulated skull and mandible, atlas-axis and first cervical vertebrae. From the upper part of the Los Molles Formation, Cuyo Group, Emileia giebeli and Emileia multiformis subzone.
Marmornectes
Marmornectes is a pliosaur with a long and narrow snout from the Oxford Clay. It was described and named by Ketchum and Benson (2011a). The type and only specimen (BEDFM 1999.201) comprises a substantially complete skeleton including a partial skull. Marmornectes is similar to Peloneustes but possesses some basal characters that were lost in Peloneustes […]
Megalneusaurus
Under construction
Meyerasaurus
M. victor was originally described and figured by Fraas (1910). Historically, M. victor has been regarded as a species of Rhomaleosaurus and is often associated in the literature under the defunct name ‘Thaumatosaurus’. See my article about this history of ‘Thaumatosaurus‘ here. Smith and Vincent (2010) identified M. victor as generically distinct from Rhomaleosaurus and erected […]
Monquirasaurus
Monquirasaurus is a giant pliosaurid from Colombia, South America. Originally named by Hampe (1992) as a species of Kronosaurus (K. boyacensis), the skeleton was later allocated to the new genus Monquirasaurus by Noè and Gómez-Pérez (2021). The genus name derives from ‘Monquira’, the administrative division in which the holotype was discovered.
Pachycostasaurus
Pachycostasaurus is approximately 3.1 meters long. Its rib cage and vertebrae exhibit thickened bone (Cruickshank et al. 1996) a condition termed pachyostosis. This heavy ossification is unusual in plesiosaurs (another exception may include Kronosaurus), although it is common in basal sauropterygians, especially pachypleurosaurs. Pachycostasaurus probably used the extra ballast provided by its heavy bones to […]
Peloneustes
Peloneustes is one of several pliosaur genera from the Oxford Clay Formation in the UK. It is one of the better known pliosaurs from this horizon, represented by 12 nearly complete skulls and several skeletons. It is also the most abundant pliosaur from the Peterborough Member of the Oxford Clay Formation (Ketchum and Benson 2011b). […]
Rhomaleosaurus
Rhomaleosaurus is the largest known Lower Jurassic pliosaur and was the top predator in early Jurassic marine ecosystems. It has a reinforced skull to help resist torsion and a ferocious set of teeth, a combination of characters perfect for snatching and killing cephalopods, fish, and other marine reptiles. Historically, the genus Rhomaleosaurus has been interchangeable […]
Simolestes
Simolestes has variously been allied with the Pliosauridae and the Rhomaleosauridae. The most noticeable difference between Simolestes and the other pliosaur taxa from the Oxford Clay (Liopleurodon, Peloneustes, Pachycostasaurus), is its much shorter snout and mandibular symphysis, a character is shares with the Rhomaleosauridae. However, this is probably a convergent character. Older descriptions of this […]
Stenorhynchosaurus
Under construction
Thalassiodracon
The species Plesiosaurus hawkinsii was introduced in 1838 for a small plesiosaurian from Street, Somerset. The new genus name Thalassiodracon was erected decades later following an examination of a referred skull in Cambridge (CAMSM J.46986). Thalassiodracon is from the Rhaetian-Hettangian boundary, so it is latest Triassic or earliest Jurassic in age. Thalassiodracon means ‘Sea Dragon’, […]
Thaumatodracon
Thaumatodracon is a relatively large rhomaleosaurid from the Lower Jurassic (Sinemurian) of Lyme Bay – the coast between Lyme Regis and Charmouth – UK. The holotype specimen (NLMH 106.058) is an almost complete skull and cervical (neck) series. It has a 60 cm long skull, and based on comparison with other rhomaleosaurids I estimate its total […]